Australopithecus

replica skull of Lucy

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Australopithecus, (Latin: “southern ape”) (genus Australopithecus), group of extinct primates closely related to, if not actually ancestors of, modern human beings and known from a series of fossils found at numerous sites in eastern, north-central, and southern Africa. The various species of Australopithecus lived 4.4 million to 1.4 million years ago (mya), during the Pliocene and Pleistocene epochs (which lasted from 5.3 million to 11,700 years ago). The genus name, meaning “southern ape,” refers to the first fossils found, which were discovered in South Africa. Perhaps the most famous specimen of Australopithecus is “Lucy,” a remarkably preserved fossilized skeleton from Ethiopia that has been dated to 3.2 mya.

As characterized by the fossil evidence, members of Australopithecus bore a combination of humanlike and apelike traits. They were similar to modern humans in that they were bipedal (that is, they walked on two legs), but, like apes, they had small brains. Their canine teeth were smaller than those found in apes, and their cheek teeth were larger than those of modern humans.

hominid fossil sites in sub-Saharan Africa

The general term australopith (or australopithecine) is used informally to refer to members of the genus Australopithecus. Australopithecines include the genus Paranthropus (2.3–1.2 mya), which comprises three species of australopiths—collectively called the “robusts” because of their very large cheek teeth set in massive jaws. Non-australopithecine members of the human lineage (hominins) include Sahelanthropus tchadensis (7–6 mya), Orrorin tugenensis (6 mya), Ardipithecus kadabba (5.8–5.2 mya), and Ar. ramidus (5.8–4.4 mya)—that is, pre-Australopithecus species that are considered to be ancient humans—and one additional species of early human, Kenyanthropus platyops (3.5 mya). The first undisputed evidence of the genus Homo—the genus that includes modern human beings—appears as early as 2.8 mya, and some of the characteristics of Homo resemble those of earlier species of Australopithecus; however, considerable debate surrounds the identity of the earliest species of Homo. In contrast, remains older than six million years are widely regarded to be those of fossil apes.